If all else fails, immortality can always be assured by spectacular error.
—John Kenneth Galbraith
Many people have the tendency to take some characters and narrowly define them based on one action or event, often to the exclusion of other, often more important, actions of that character.
Often, this is perfectly understandable, if that action or event stands out more than anything else the character did. But just as often it doesn't, and it can seem as if some people just haven't been paying attention.
Let's take a hypothetical show Bob Loves Alice. There is a fan favorite episode where Bob gets caught up in a game of darts. He isn't that good, and he never really tries darts again. A bunch of fans like that episode enough that, in loads of Fan Fiction, Bob sucking at darts is his defining characteristic and it's what everyone knows him for, when in actual Canon, it's never really mentioned again.
On the internet, it is usually the person's never live it down-ed aspect that undergoes Memetic Mutation.
This can also turn even worse when those fans start Running the Asylum in that show's later seasons, and now everyone on the show talks about how Bob sucks at darts, while ignoring all the other things Bob has done since then. But you never see Bob playing darts, because surely we've seen enough of that. No, we haven't. We've seen it once. And since this is placing all of Bob's more prominent traits and events behind this minor one, it even risks taking away a dimension of Character Depth.
When the show decides to remind you that Bob-The-Sucky-Dart-Player is not his canon personality, that's Forgot Flanders Could Do That.
Remember When You Blew Up a Sun? seems like this, but it merely harks back to a character's Crowning Moment of Awesome, not pretend that's all the character does. Also related is One-Scene Wonder, where a small guest star starts to dominate Fanfic.
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