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''I/O'', like the ''Infinity'' series, has multiple protagonists, in this case four. Route A features Hinata Aoi, an [[Ordinary High School Student]] who has become depressed after the disappearance of his twin sister, Mutsuki. Route B features Ishtar, a young woman in charge of a hacking group known as Criminal. Route C also features Ishtar, but she seems different from herself in Route B. Route D features "He", a mysterious man who is a legend on the Internet. Once all four of their routes have been completed, they unlock Route E, the true route.
The game was originally released on the [[
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* [[Ambiguous Gender]]- Enlil, who is never specifically identified as a man or a woman. {{spoiler|Route E finally confirms that Enlil is a woman.}}
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