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The book follows two foreign princes, Pyrocles and Musidorus, who arrive in Arcadia and in order to get close to two princess, Philoclea and Pamela respectively, they disguise themselves. Pyrocles [[Gender Bender|dresses as a woman]] and takes on the imaginative name Cleophila (changed to Zelmane in the New Arcadia), while Musidorus dresses as a humble shepherd and starts calling himself Dorus. They engage in various adventures and also have to deceive the Duke of Arcadia, Basilius, and his wife, Gynecia. After each book and before the beginning of the next are selections of poems called the Eclogues, which are sung by various characters, often the Arcadian shepherds [[Hero of Another Story|Strephon and Klaius]].
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