Dinosaurs (useful notes): Difference between revisions

 
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{{tropeUseful Notes}}
[[Blatant Lies|A brief summary]] of [[Tropesaurus Index|dinosaurs]], for those of us who wish to address a few problems where [[Somewhere a Palaeontologist Is Crying]].
 
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* [http://albertonykus.deviantart.com/art/Why-Birds-are-Dinosaurs-193639479 Birds are dinosaurs.] Because there are still birds alive today, dinosaurs are [[Science Marches On|technically not exinct]]!
 
All clear? Right. On to the next topic.
 
== The Dinosaur Family Tree ==
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Dinosaurs are divided into two different groups, and those two groups are ''not'' "meat-eaters" and "plant-eaters". In fact, some plant-eaters were more closely related to the meat-eaters than they were to other plant-eaters. In any case, this kind of grouping fails to account for those dinosaurs that were omnivorous, insectivorous, piscivorous etc., and is largely frowned upon within phylogenetic biology in any case, since in evolution a species can switch from one form of diet to another at the drop of a geological hat.
 
The first group is known as the Saurischia, or “lizard-hipped” dinosaurs, which include all the meat-eating dinosaurs, or theropods, and those long-necked, plant-eating dinosaurs, the sauropodomorphs (which include the giant sauropods). Despite how vividly different a ''[[Tyrannosaurus Rex|Tyrannosaurus]]'' looks to a ''[[Stock Dinosaurs True Dinosaurs|Brachiosaurus]]'', the earliest theropods and sauropods looked very similar. One early saurischian, ''[[Useful Notes/Prehistoric Life /Dinosaurs|Eoraptor]]'', has been considered a theropod, a sauropodomorph, or a basal saurischian that didn’t belong to either group.
 
The second group is known as the Ornithischia, or “bird-hipped” dinosaurs, and are mostly herbivorous dinosaurs, very different in body-shape both when compared with saurischians and when compared with each other. This group includes the ornithopods, such as the duck-billed hadrosaurs and ''[[Stock Dinosaurs True Dinosaurs|Iguanodon]]'', the thyreophorans, such as the stegosaurs and ankylosaurs, and the marginocephalians, such as the thick-skulled pachycephalosaurs and horned and frilled ceratopsians.
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When the names of the Saurischia and the Ornithischia were first coined, they were differentiated chiefly by their pelvic arrangement, with the saurischian hip bones arranged more like a lizard hip, the pubis bone pointing forwards (but this is the case in crocs, mammals and turtles as well, so it is not something unique about lizards), and the ornithischian hip bones more like an avian hip, the pubis bone pointing backwards. This latter was possibly an adaptation designed to accomodate longer guts for the digestion of vegetation, which is notoriously hard to digest.
 
That much most textbooks and children's books will tell you, but the curious thing is that birds are theropod saurischians, not ornithischians, even though you'd expect some sort of bird-like hip arrangement to appear in their ancestors. So how did theropods bring about the bird group if they had the lizard hip, not the bird hip? Fortunately, this is quickly resolved: ornithischians are not the only dinosaurs to have bird-like hips. Many maniraptorans, those theropod dinosaurs which include birds as well as the theropods most closely related to birds (and are therefore what most people refer to when they talk about bird-like dinosaurs), have a bird-like hip arrangement, which appears to be a side effect of some adopting a knee-driven running style, and of others being weird herbivorous theropods. Interestingly, the most basal ornithischians, such as ''[[Useful Notes/Prehistoric Life /Dinosaurs|Pisanosaurus]]'', hadn't yet evolved the backwards-pointing pubis.
 
At first, you might think this rather puts a hole through our lovely little classification. How can a saurischian be a saurischian if it's got bird-like hips, a characteristic of ornithischians? The pelvic arrangement is not the only way to differentiate between the two groups. Luckily, ornithischians can still be kept separate from saurischians by another diagnostic: all ornithischians have a U-shaped protruding bone at the front of the lower jaw called a predentary bone, often ending with a beak, and most of them have a row of chewing teeth lined up either side of the face, giving it a certain "hollow cheek" look like that of a horse. This suggests that many ornithischians chewed their food, unlike saurischians. Also, saurischians have air sacs in the vertebrae that ornithischians lacked, air sacs which the birds inherited, and which help to keep birds lightweight when flying.
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Birds, considered as a distinct class of vertebrate in traditional systematics, are actually theropod dinosaurs, a hypothesis already proposed in the eighteenth century after the discovery of the famous ''[[Stock Dinosaurs True Dinosaurs|Archaeopteryx]]'' (an animal with dinosaurian skeleton ''and'' feathered wings and tail) but rejected by most scientists for a long amount of time. The link between dinosaurs and birds through ''Archaeopteryx'' was resurrected again in the 1960s, and has been definitively proven only in the 1990's by the long list of feathered dinosaurs and early birds recently found in the fossil record, which show strong anatomical similarities.
 
Impressions of protofeathers and true feathers in the fossils make for good evidence, and blend into each other so seamlessly that telling apart bird-like dinosaurs (such as ''[[Stock Dinosaurs True Dinosaurs|Velociraptor]]'', ''[[Useful Notes/Prehistoric Life /Dinosaurs|Caudipteryx]]'', ''[[Useful Notes/Prehistoric Life /Dinosaurs|Beipiaosaurus]]'', ''[[Useful Notes/Prehistoric Life /Dinosaurs|Sinosauropteryx]]'') and dino-like birds (''[[Useful Notes/Prehistoric Life /Other Extinct Creatures|Jeholornis]]'', ''[[Useful Notes/Prehistoric Life /Other Extinct Creatures|Confuciusornis]]'' and so on) has become very difficult today.
 
Interestingly, some ornithischian dinosaurs like ''[[Useful Notes/Prehistoric Life /Dinosaurs|Psittacosaurus]]'' and ''[[Useful Notes/Prehistoric Life /Dinosaurs|Tianyulong]]'' have been discovered with quills or other structures that strongly resemble feathers, which has raised intriguing questions about what dinosaurs looked like. Some think all dinosaurs originally had some sort of covering at the start of their evolution; then this covering (perhaps some sort of hypothetical "down") was lost in some lineages, or at least modified into other specialized structures (the quills on ''Psittacosaurus'' or even the dorsal spines on the sauropod ''[[Stock Dinosaurs True Dinosaurs|Diplodocus]]'' may have this origin). The fact that the closely-related pterosaurs have a covering made of a sort of hollow hair seems to confirm this hypothesis. All the same, it does not eliminate the chance that the pterosaurs stumbled across the same solution on their own. If so, it would have been an example of convergent evolution.
 
Main points:
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And now, the history. We've got a lot to cover - ninety thousand times more dinosaur history than human history, for a start.
 
Dinosaurs dominated all the land environments of the Mesozoic era, the era which covers a vast geological timescale from 251 to 65.5 million years ago. And, unlike what many movies and illustrations show, not all dinosaurs lived at the same time. The earliest dinosaur forms, such as ''[[Useful Notes/Prehistoric Life /Dinosaurs|Eoraptor]]'' and ''[[Useful Notes/Prehistoric Life /Dinosaurs|Eocursor]]'', appeared during the middle of the Triassic period, the first of the three geological periods which make up the Mesozoic era, and for most of the Triassic period they were background detail.
 
The Triassic wasn't a pleasant time for life - they were living in the wake of the greatest mass extinction of the Phanerozoic eon (the last five hundred million years, roughly speaking), which had wiped out almost 95% of all living species. The continents of the world had fused into one supercontinent, a giant landmass called Pangea, and vast deserts covered the innermost areas. (thisThis could be one reason why the Permian-Triassic mass extinction was so devastating; [http://www.nature.com/articles/srep43630 another hypothesis] is that it happened when massive eruptions produced lots of volcanic gases and led to a short ice age: cold used to cause large extinctions all on its own, even without acidic fallout on top (how this happened depends on how all those events are dated). The surviving animals had to recover pretty quickly, and many animal classes battled it out.
 
The mammal-like "reptiles", which had ruled in waves during the Permian period, were the first to spread on the planet, and it looked like they were set to rule it all again. There were two groups of these, mostly the herbivorous dicynodonts and the cynodonts. Mammals evolved from the latter set; the former died out without leaving any descendants.
 
However, both groups quickly found themselves under stiff competition against the first archosaur reptiles, from which the dinosaurs would emerge. The archosaurs were particularly effective thanks to their water-conserving adaptations, which were typical of most reptiles. Quickly, the archosaurs divided into several groups, such as rauisuchians, aetosaurs, phytosaurs and proterosuchids, and by seizing control they reduced the variety of the mammal-like reptiles, which survived only through their smaller species. Later, cynodonts evolved many special features that are typical of modern mammals today, humans included.
 
In the midst of this inter-group conflict, roughly 230 million years ago, the early dinosaurs appeared. They were descended from tiny archosaurs such as the 1 ft-long ''[[Useful Notes/Prehistoric Life /Non Dinosaurian Reptiles|Lagosuchus]]'', but they did not make their impact felt until ten million years later, when they grew from small, unassuming bipeds to impressively large forms. The three main lineages were forged at this crucial time: the meat-eating theropods, the long-necked sauropodomorphs, and the plant-eating ornithischians, although at this stage they all looked like variations of the thin, elegant Theropods like ''[[Stock Dinosaurs True Dinosaurs|Coelophysis]]'' and its relatives, and it's quite possible that all of them were omnivorous initially.
 
They were notable for taking the bipedal stance, which was also adopted by some triassic archosaurs related to modern crocodilians, such as ''[[Useful Notes/Prehistoric Life /Non Dinosaurian Reptiles|Ornithosuchus]]'' and the deceptively-dinosaurian ''[[Useful Notes/Prehistoric Life /Non Dinosaurian Reptiles|Effigia]]''. Indeed, some of these were mistaken for dinosaurs when they were first discovered. To walk in the upright stance, early dinosaurs developed a horizontal backbone but vertical joints to the pelvis, which meant that their legs were tucked underneath their body to support their weight. This also permitted them an exceptionally good turn of speed, and with their long tails to serve as counterbalances, the dinosaurs had hit upon a good design feature which would serve them well again and again.
 
Even with all the similarly-adapted archosaurs, Dinosaurs were notably successful at this early stage, and by the end of the Triassic period they had diversified into some of the largest animals ever to appear on the land. Early sauropodomorphs in particular (traditionally called "prosauropods", which means "before the sauropods") reached lengths and heights never seen before, like ''[[Stock Dinosaurs True Dinosaurs|Plateosaurus]]'', which could grow up to twenty feet long, and the even larger ''[[Useful Notes/Prehistoric Life /Dinosaurs|Riojasaurus]]'' and ''[[Useful Notes/Prehistoric Life /Dinosaurs|Melanorosaurus]]'', both elephant-sized and thirty feet long. On the other hand, predatory theropods remained generally small in the Triassic, with some exceptions such as ''[[Godzilla|Gojira]][[Useful Notes/Prehistoric Life /Dinosaurs|saurus]]'' (whoever said that paleontologists had no sense of humour?), which could reach 15 ft in length.
 
Most ornithischians were still small plant-eaters, though even here, specialized new forms were emerging - such as the heterodontosaurids, creatures with large canine teeth that may have been used in mating disputes. As an aside, ''Heterodontosaurus'' is often speculated to be the male form of another heterodontosaurid species, and its tusks were thought to be weapons used against other males in mating disputes, rather like the tusks used by musk deer today. They are almost never talked about in pop culture, though the scientific interest for them sky-rocketed with the discovery of proto-feathers on the fossils of ''Tianyulong''. No one knows what became of the heterodontosaurids after their heyday in the Triassic and Early Jurassic periods, though the species ''Echinodon'', which lived during the Early Cretaceous period, implies that they may have survived quietly for millions of years long after the Jurassic.
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The purpose of these plates is unclear, but the purpose of their tail spikes and shoulder spikes was arguably for self-defence against any animal that tried to hurt or kill the animal. Their cousins, the ankylosaurs, took the scutes and developed them into stronger armour all along their backs and even, in some cases, along their undersides. The species ''Ankylosaurus'' even had armour-plated eyelids, and a thickened block of bony tissue at the end of their tails to act as clubs in case the armour wasn't a good enough hint for some carnivores. They came into greater prominence during the Cretaceous period, when the stegosaurs died out.
 
Besides the thyreophorans, there were the cerapods - the collective name for both the ornithopods and the marginocephalians. Ornithopods became more diverse during the Jurassic (''[[Useful Notes/Prehistoric Life /Dinosaurs|Dryosaurus]]'' and ''[[Useful Notes/Prehistoric Life /Dinosaurs|Camptosaurus]]'' were widespread at that time), but they really came into their own during the Cretaceous period, possibly because of the spread of the newly-evolved flowering plants which had appeared not long before, or because they coped better with the changing climate as sea levels rose worldwide. The marginocephalians possibly had Late Jurassic roots, but like the ornithopods they are better known for their Cretaceous forms, and originally evolved from very small ornithopod-like forms such as ''[[Useful Notes/Prehistoric Life /Dinosaurs|Yinlong]]''.
 
Main points:
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Dinosaurs continued to diversify further in the Cetaceous, but times were changing and things weren't what they used to be.
 
Among ornithischians, marginocephalians differentiated into two subgroups. The pachycephalosaurs remained small and bipedal but developed thick skulls for uncertain purposes, while the ceratopsians became progressively heavier, and quadrupedal. Their beaks became parrot-shaped, they evolved protrusions from their cheeks and a large "frill" from the backs of their heads, and some of the later ones had impressive sets of horns and spikes. The earliest ceratopsians, if you don't count creatures like ''Psittacosaurus'', were the protoceratopsids. Later, these smallish (by dinosaur standards - by modern standards, some would be quite respectable in size) animals evolved into the large, rhino-resembling ceratopsids, including ''[[Useful Notes/Prehistoric Life /Dinosaurs|Chasmosaurus]]'', ''[[Stock Dinosaurs True Dinosaurs|Styracosaurus]]'', the ever-popular ''[[Stock Dinosaurs True Dinosaurs|Triceratops]]'', and ''[[Useful Notes/Prehistoric Life /Dinosaurs|Torosaurus]]'', though recently there have been discussions about whether or not ''Torosaurus'' and ''Triceratops'' are actually one species.
 
Strictly speaking, the split between pachycephalosaurs and ceratopsians is largely believed to have occurred in the Jurassic period, but fossils of Jurassic pachycephalosaurs have not yet been found. It is a bit of a mystery what the thick skulls of the pachycephalosaurs were used for. The impulse to say that they were used to defend the animal from predatory attack should not be indulged - the smaller creatures tend to have flatter skulls, admittedly, but the larger creatures have heads like bowling balls, which means that they have a very small surface area available for actually hitting anything. Try running into somebody while holding a bowling ball out in front of you at arm's length and you'll get some idea of how tricky this actually would be to pull off. Paleobiologists thesedays tend to believe that the skulls were used to butt the sides of rival pachycephalosaurs, perhaps over territorial or mating rights. Some pachycephalosaur skulls were surrounded by spikes and knobs of bone, and probably would have looked frightening in the eyes of a small predator or a rival, and an interesting little debate is going on over whether the spikier forms, like ''Stygimoloch'' and ''Dracorex'', are really different forms of the less spikier species - they may be child forms, or male-female forms, for example.
 
Ornithopods, meanwhile, included both small species such as ''[[Useful Notes/Prehistoric Life /Dinosaurs|Hypsilophodon]]'' and huge species, the most spectacular of which were the styracosternans. These included the hadrosaurs, giant four-legged beasts with duck-like bills for cropping plants over a wide area, and iguanodontids like ''Iguanodon'', all of which had a specialized front foot with a thumb-spike, three padded fingers to support the animal's weight during quadrupedal locomotion, and a flexible little finger for grasping food. The hadrosaurs do not have the thumb spike, and it was probably used for decoration or feeding rather than for defence. Between these two extremes were middle-sized creatures like ''[[Useful Notes/Prehistoric Life /Dinosaurs|Tenontosaurus]]'' and ''[[Useful Notes/Prehistoric Life /Dinosaurs|Muttaburrasaurus]]''.
 
Hadrosaurs were the more successful of the styracosternans, and came with a broad variety of crests and nasal passages decorating the skulls, which were probably used to tell each other apart in case one hadrosaur made the mistake of trying to mate with another hadrosaur from a different species. The styracosternans as a whole were successful, perhaps because of their chewing teeth, which could pulp up even the toughest plant material. Fossil nests and eggs show that hadrosaurs were devoted mothers, like many modern birds are, and looked after their babies until they were old enough to fend for themselves. ''Maiasaura'' is probably the best known hadrosaur whose fossils include evidence of maternal care - indeed, ''Maiasaura'' means "good mother lizard".
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During their time on Earth, the dinosaurs thrived in diverse terrestrial habitats, from swampy terrain and dense forests to open prairies and the driest of deserts. Some even weathered the harsh winter conditions of Antarctica and Australia (which were near the South Pole at the time). However, the dinosaurs did not survive the Cretaceous-Paleogene (formerly known as the Cretaceous-Tertiary) mass extinction event which wiped out roughly 65% of all living things, at least according to the fossil record. This is that eponymous event which underscores the trope [[Phlebotinum Killed the Dinosaurs]], and if you read some of the hypotheses about how they died, they often run a lot like that trope.
 
There have been several hypotheses in the past about this event, but most of them are bunk and the rest are on shaky ground at best. In the early days of palaeontology, when snobbery of the past was widespread and extinction more or less meant you were an inferior species, it was believed that the dinosaurs simply became [[Too Dumb to Live]], or to put it more scientifically, their craniums housed brains which were inefficient by dint of being too small, and so they were outwitted by the smaller but much cleverer mammals. Some have suggested that egg-devouring mammals were responsible, but there is no evidence that the mammals consumed eggs, at least not in bulk. Some suggested that the dinosaurs found it harder to supply their large bodies with oxygen, ignoring the fact that Late Cretaceous dinosaurs were pretty tame compared with the giganic sauropods which had dominated in the Jurassic (and which, as far as can be made out, never had breathing difficulties). Some have suggested that a nearby supernova caused cosmic rays to penetrate the atmosphere and destroy the dinosaurs, but a supernova that close to the Earth would have done considerably more than wipe out a few little reptiles on its surface, and in any case the claim is not justified by evidence.
 
Here is the most likely hypothesis currently available. The extinction began with an increase in volcanic activity during the last few million years of the Cretaceous period, which would have introduced toxic gases and ash clouds into the atmosphere. This interfered with the relatively stable weather conditions the dinosaurs had enjoyed all over the globe (most of the Mesozoic era was comparatively stable, at least when compared with the turbulent climate changes of most of the Cenozoic era). Certainly, the fossil evidence suggests they were already in a state of decline (at least in North America) when the fateful meteor, about 65.5 million years ago, [[Rocks Fall, Everyone Dies|collided with the Gulf of Mexico]], producing the Chicxulub crater.
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* Not all the surviving lineages are still alive today. Multituberculate mammals and champsosaurs survived the extinction but died later in the Cenozoic.
 
[[Vindicated by History|Dinosaurs]] are now a pretty big hit in popular culture. Just go see the [[Tropesaurus Index]], and you'll find links to the legacy that the dinosaurs left behind. Yes, that includes the bird index, too.
 
If you are interested in specific kinds of dinosaurs, just [[Stock Dinosaurs True Dinosaurs|check]] [[Useful Notes/Prehistoric Life /Dinosaurs|here]] some info.
 
{{reflist}}
[[Category:Useful Notes]]
[[Category:Tropesaurus Index]]
[[Category:Dinosaurs]]